Research/Areas of Interest:

I no longer maintain an active research laboratory and do not accept dissertation students.

I remain fully engaged in graduate education through my roles in the Immunology Program and Tufts PREP.

My research most recently focused on the longstanding observation that as people age they often become more susceptible to infectious diseases and how a changing immune system contributes to this phenomenon.


  • Doctor of Medicine, Albert Einstein College of Medicine, New York, United States, 1960
  • Bachelor of Science, University of Wisconsin Colleges, Madison, United States, 1956